古脊椎动物学报

ZHANG Zhao-Qun YANG RuiKey Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Scicences Beijing 100044 University of Chinese Academy of Sciences Beijing 100049

WU Wen-Yu MENG Jin YE Jie NI Xi-Jun BI Shun-DongKey Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing 100044 China Division of Paleontology American Museum of Natural -History New York NY 10024 USA Department of Biology lndiana University of Pennsylvania Indiana PA 15705 USA

WANG Shi-Qi DENG TaoKey Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing 100044 CAS Center for Excellence in Tibetan Plateau Earth Sciences Beijing 100101 Key Laboratory of Economic Stratigraphy and Palaeogeography Chinese Academy of Sciences (Nanjing Institute of Geology and Palaeontology) Nanjing 210008

XU Shi-Chao YOU Hai-Lu WANG Jia-Wei WANG Suo-Zhu YI Jian JIA LeiShanxiMuseum of Geology Taiyuan 030024 China Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing 100044 China Faculty of Geology Lomonosov Moscow State University Moscow 119991 Russian Federation